Paris Saint-Germain players can ill afford to cruise through the season as coach Luis Enrique warned he expected them to play out of their comfort zone.

Rennes, four days after firing coach Habib Beye, stunned Paris Saint-Germain 3-1 on Friday, snapping the defending champions' seven-match winning run in Ligue 1.

Liverpool beat off competition from Chelsea to sign Rennes defender Jeremy Jacquet for a reported fee of up to £60 million ($82 million) on Monday.

A youthful Liverpool side lost 2-1 at Union Saint-Gilloise in their final Europa League group game on Thursday, while Villarreal snatched a place in the last 16 from Rennes with a dramatic victory in France.

Kylian Mbappe made his return from a post-World Cup holiday but Paris Saint-Germain went down 1-0 away to Rennes in Ligue 1 on Sunday, a result which leaves them just three points clear at the top of the table halfway through the season.